Cyclosorus ciliatus (Wall. ex Benth.) Panigrahi

Family

Thelypteridaceae

Nomenclature

Cyclosorus ciliatus (Wall. ex Benth.) Panigrahi, Res. J. Pl. Environ. 9: 66. 1993. – Aspidium ciliatum Wall. ex Benth., Fl. Hongk.: 455. 1861. – Lastrea calcarata var. ciliata (Wall. ex Benth.) Bedd., Handb. Ferns Brit. India: 235, f. 121. 1883. – Thelypteris ciliata (Wall. ex Benth.) Ching, Bull. Fan Mem. Inst. Biol. 6: 289. 1936; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 362. 1941; Holttum, Rev. Fl. Malaya ed. 1, 2:250, f. 142. 1955 [‘1954’]; Holttum, Dansk Bot. Ark. 20: 22. 1961; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 78. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 68. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 401, f. 39. 6–9. 1988; Newman et al., Checkl. Vasc. Pl. Lao PDR: 33. 2007. – Trigonospora ciliata (Wall. ex Benth.) Holttum, Blumea 19: 29. 1971; Reinwardtia 8: 507. 1974; Holttum, Fl. Males., Ser. II, Pterid. 1: 375, f. 5c–e. 1982 [‘1981’]; Boonkerd & Pollawatn, Pterid. Thailand: 231. 2000. – Type: "Bowman" (Bowring), Hong Kong (K, lectotype (Holttum 1974)) .

Description

Rhizome short, erect; scales up to 3 by 1.5 mm, membranous, appressed, brown, hairy. Stipes stramineous to greyish, hairy throughout, up to 30 cm long. Soriferous fronds usually taller and larger than sterile ones, oblong-lanceolate, acute at apex, 13–35 by 6–13 cm; lowest pinnae shortened, deflexed, pinnae of lower 1/3 largest, patent, subsessile, linear-lanceolate, caudately acuminate at apex, broadly cuneate at base, often auricled; rachis and costa densely hirsute; segments falcate, oblique, rounded at apex, subentire, hairy at margin; chartaceous or thicker, deep green; veins pinnate, veinlets simple, basal anterior ones running to sinus, hairy. Sori subcostular, often confluent at maturity; indusia firm, persistent, hairy .

Distribution in Thailand

NORTHERN: Chiang Mai, Chiang Rai, Phrae, Tak, Phitsanulok; NORTH-EASTERN: Phetchabun, Loei; SOUTH-EASTERN: Chanthaburi; PENINSULAR: Nakhon Si Thammarat, Trang, Yala.

Distribution in Laos

Luang Phrabang.

Wider Distribution

Nepal to W Malesia, north to Hong Kong.

Ecology

Usually on muddy stream-beds in dense forests at 500–1700 m alt.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Mitsuta et al. T-50256, Chiang Mai (QBG); Middleton et al. 4927, Chiang Mai, Doi Inthanon National Park (E); Middleton et al. 4652, Chanthaburi, Namtok Phlio National Park (E).

Habit

Habit

Habit

Habit

Habit

Habit

Rhizome

Rhizome

Crozier

Crozier

Stipe and stipe scales

Stipe and stipe scales

Lamina

Lamina

Upper surface of lamina

Upper surface of lamina

Basal pinnae

Basal pinnae

Venation

Venation

Frond apex with older sori

Frond apex with older sori

Lower surface of lamina

Lower surface of lamina

Pinnae with young sori

Pinnae with young sori

Young sori

Young sori

Young sori

Young sori

Older sori

Older sori

Site hosted by the Royal Botanic Garden Edinburgh. Content managed by Stuart Lindsay, Gardens by the Bay, Singapore and David Middleton, Singapore Botanic Gardens. Last updated 16 November 2016.