Ferns of Thailand, Laos and Cambodia

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Family

Thelypteridaceae

Nomenclature

Cyclosorus nudatus (Roxb.) B.K.Nayar & S.Kaur, Companion Beddome's Handb. Ferns Brit. India 66, 69. 1974. – Polypodium nudatum Roxb., Calcutta J. Nat. Hist. 4: 491. 1844. – Pronephrium nudatum (Roxb.) Holttum, Blumea 20: 111. 1972; Boonkerd & Pollawatn, Pterid. Thailand: 166, 167, 227. 2000. – Thelypteris nudata (Roxb.) C.V.Morton, Contr. U.S. Natl. Herb. 38: 352. 1974; Tagawa & K.Iwats., Acta Phytotax. Geobot. 26: 170. 1975; Tagawa & K.Iwats., Fl. Thailand 3: 411, f. 41.1, 42.2–3. 1988.

Nephrodium moulmeinense Bedd., Suppl. Ferns Brit. Ind.: 18. 1876; Bedd., Handb. Ferns Brit. India: 275.f. 141. 1883. – Dryopteris moulmeinense (Bedd.) C.Chr., Index Filic.: 278. 1905; Contr. U.S. Natl. Herb. 26: 330. 1931.

Polypodium multilineatum Wall. ex Hook., Sp. Fil. 5: 11. 1863. – Nephrodium multilineatum (Wall. ex Hook.) Bedd., Suppl. Ferns Brit. Ind.: 80. 1892, nom. illeg. – Abacopteris multilineata (Wall. ex Hook.) Ching, Bull. Fan Mem. Inst. Biol. 8: 253. 1938; Holttum, Rev. Fl. Malaya ed. 1, 2: 297. 1955 [‘1954’]; Holttum, Dansk Bot. Ark. 20: 23. 1961. – Cyclosorus multilineatus (Wall. ex Hook.) Tardieu & C.Chr., Fl. Indo-Chine 7(2): 358. 1941. – Thelypteris multilineata (Wall. ex Hook.) Morton, Amer. Fern J. 49: 113. 1959; K.Iwats., J. Jap. Bot. 38: 315. 1963; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 80. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 70. 1967.

Dryopteris urophyllum auct. non (Mett.) C.Chr.: Bonap., Notes Ptérid. 14: 49. 1923. – Nephrodium urophyllum auct. non (Mett.) Keys.: E.Smith, J. Siam. Soc. Nat. Hist. Suppl. 8: 5. 1929.

Description

Rhizome creeping, about 5 mm diam.; scales caducous, dark brown, hairy. Stipes 35–80 cm long, scaly at base. Laminae oblong, 50–100 by 25–50 cm; lateral pinnae lanceolate, sessile, ascending, gradually narrowing towards long-acuminate apex, rounded to narrowly cuneate at base, subentire or crenate, 18–40 by 2.5–5 cm; terminal pinna like lateral ones, rounded to subtruncate at base; marginal lobes acute at apex, with cartilaginous margin; chartaceous, green, verrucose on lower surface; venation meniscioid. Sori rather close to excurrent veinlets or medial in two rows between costules; indusia setose; sporangia glabrous .

Distribution in Thailand

NORTHERN: Mae Hong Son, Chiang Mai, Chiang Rai, Nan, Lampang, Phrae, Tak, Phitsanulok; SOUTH-WESTERN: Kanchanaburi; CENTRAL: Nakhon Nayok; SOUTH-EASTERN: Chanthaburi, Trat; PENINSULAR: Surat Thani, Phangnga, Nakhon Si Thammarat.

Distribution in Cambodia

Mondulkiri.

Wider Distribution

Himalaya, Burma, China and N Vietnam.

Ecology

On rather dry ground in light shade or in dense forests at medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Cambodia

Long et al. CL466, Mondulkiri (P).

	Ferns of Thailand, Laos and Cambodia
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Cyclosorus nudatus (Roxb.) B.K.Nayar & S.Kaur Warning: count(): Parameter must be an array or an object that implements Countable in C:\Data\serverroots\htdocs\ThaiFerns\factsheets\functions.php on line 17 Family Thelypteridaceae Nomenclature Cyclosorus nudatus (Roxb.) B.K.Nayar & S.Kaur, Companion Beddome's Handb. Ferns Brit. India 66, 69. 1974. – Polypodium nudatum Roxb., Calcutta J. Nat. Hist. 4: 491. 1844. – Pronephrium nudatum (Roxb.) Holttum, Blumea 20: 111. 1972; Boonkerd & Pollawatn, Pterid. Thailand: 166, 167, 227. 2000. – Thelypteris nudata (Roxb.) C.V.Morton, Contr. U.S. Natl. Herb. 38: 352. 1974; Tagawa & K.Iwats., Acta Phytotax. Geobot. 26: 170. 1975; Tagawa & K.Iwats., Fl. Thailand 3: 411, f. 41.1, 42.2–3. 1988. Nephrodium moulmeinense Bedd., Suppl. Ferns Brit. Ind.: 18. 1876; Bedd., Handb. Ferns Brit. India: 275.f. 141. 1883. – Dryopteris moulmeinense (Bedd.) C.Chr., Index Filic.: 278. 1905; Contr. U.S. Natl. Herb. 26: 330. 1931. Polypodium multilineatum Wall. ex Hook., Sp. Fil. 5: 11. 1863. – Nephrodium multilineatum (Wall. ex Hook.) Bedd., Suppl. Ferns Brit. Ind.: 80. 1892, nom. illeg. – Abacopteris multilineata (Wall. ex Hook.) Ching, Bull. Fan Mem. Inst. Biol. 8: 253. 1938; Holttum, Rev. Fl. Malaya ed. 1, 2: 297. 1955 [‘1954’]; Holttum, Dansk Bot. Ark. 20: 23. 1961. – Cyclosorus multilineatus (Wall. ex Hook.) Tardieu & C.Chr., Fl. Indo-Chine 7(2): 358. 1941. – Thelypteris multilineata (Wall. ex Hook.) Morton, Amer. Fern J. 49: 113. 1959; K.Iwats., J. Jap. Bot. 38: 315. 1963; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 80. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 70. 1967. Dryopteris urophyllum auct. non (Mett.) C.Chr.: Bonap., Notes Ptérid. 14: 49. 1923. – Nephrodium urophyllum auct. non (Mett.) Keys.: E.Smith, J. Siam. Soc. Nat. Hist. Suppl. 8: 5. 1929. Description Rhizome creeping, about 5 mm diam.; scales caducous, dark brown, hairy. Stipes 35–80 cm long, scaly at base. Laminae oblong, 50–100 by 25–50 cm; lateral pinnae lanceolate, sessile, ascending, gradually narrowing towards long-acuminate apex, rounded to narrowly cuneate at base, subentire or crenate, 18–40 by 2.5–5 cm; terminal pinna like lateral ones, rounded to subtruncate at base; marginal lobes acute at apex, with cartilaginous margin; chartaceous, green, verrucose on lower surface; venation meniscioid. Sori rather close to excurrent veinlets or medial in two rows between costules; indusia setose; sporangia glabrous . Distribution in Thailand NORTHERN: Mae Hong Son, Chiang Mai, Chiang Rai, Nan, Lampang, Phrae, Tak, Phitsanulok; SOUTH-WESTERN: Kanchanaburi; CENTRAL: Nakhon Nayok; SOUTH-EASTERN: Chanthaburi, Trat; PENINSULAR: Surat Thani, Phangnga, Nakhon Si Thammarat. Distribution in Cambodia Mondulkiri. Wider Distribution Himalaya, Burma, China and N Vietnam. Ecology On rather dry ground in light shade or in dense forests at medium altitudes. Proposed IUCN Conservation Assessment Least Concern (LC). This species is widespread and not under any known threat. Voucher specimens - Cambodia Long et al. CL466, Mondulkiri (P). Warning: count(): Parameter must be an array or an object that implements Countable in C:\Data\serverroots\htdocs\ThaiFerns\factsheets\functions.php on line 17
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Cyclosorus nudatus (Roxb.) B.K.Nayar & S.Kaur