Cyclosorus polycarpus (Blume) Holttum

Family

Thelypteridaceae

Nomenclature

Cyclosorus polycarpus (Blume) Holttum, Rev. Fl. Malaya ed. 1, 2: 283, f. 164. 1955 [‘1954’]. – Aspidium polycarpum Blume, Enum. Pl. Javae.: 156. 1828. – Nephrodium polycarpum (Blume) Keys., Polyp. Herb. Bunge: 7. 1873; Bedd., Suppl. Ferns Brit. Ind.: 74. 1892. – Mesochlaena polycarpa (Blume) Bedd., Suppl. Ferns Brit. Ind.: 13. 1876; Bedd., Handb. Ferns Brit. India: 199, f. 100. 1883. – Sphaerostephanos polycarpus (Blume) Copel., Univ. Calif. Publ. Bot. 16: 60. 1929; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 400, f. 33. 3–4. 1941; Holttum, Blumea 19: 40. 1971; Holttum, Kalikasan 4:56. 1975; Holttum, Fl. Males., Ser. II, Pterid. 1: 448, f. 12a–c. 1982 [‘1981’]; Boonkerd & Pollawatn, Pterid. Thailand: 168, 229. 2000. – Thelypteris polycarpa (Blume) K.Iwats., Mem. Coll. Sci. Univ. Kyoto B. 31: 32. 1964; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 78. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 69. 1967; Tagawa & K.Iwats., Acta Phytotax. Geobot. 23: 54. 1968; Tagawa & K.Iwats., Fl. Thailand 3: 416. 1988. – Type: Blume, Noesa Kambangan, Java (L).

Description

Rhizome massive, erect; scales narrow, up to 20 by 1.2 mm, hairy at margin as well as on dorsal surface, concolorous brown. Stipes 25–130 cm long, 2 cm diam., hairy, bearing reduced pinnae nearly to base. Laminae 49–118(–150) by 17–52(–60) cm, oblong-lanceolate, with 37–62 pairs of full-sized pinnae; lower pinnae suddenly reduced to butterfly-like projections and to mere auricles, middle pinnae nearly straight, ascending or patent, linear, 9.5–35 by 0.9–2.5 cm, lobed to half-way towards costa; segments oblong, oblique, rounded at apex, entire; chartaceous, green; rachis and costa pubescent; veins pinnate, veinlets more than 15 pairs, simple, lower 2.5 pairs usually uniting below sinus, hirsute on both surfaces, ending in hydathodes just inside the margin. Sori medial, elongate along veinlets, up to 1.5 by 0.2 mm, confluent at maturity; indusia oblong, dark, hairy .

Distribution in Thailand

SOUTH-WESTERN: Kanchanaburi; SOUTH-EASTERN: Chanthaburi; PENINSULAR: Surat Thani, Phangnga, Nakhon Si Thanimarat, Trang, Satun, Yala, Narathiwat.

Wider Distribution

Malesia.

Ecology

On rather dry ground in open places at low to medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Middleton et al. 4779, Kanchanaburi, Thong Pha Phum National Park (E); Smith 3048, Surat Thani, Bandon (K); Middleton et al. 5358, Satun, near Thale Ban National Park (E); Smith 1865, Yala, Betong (K).

Habit

Habit

Habit

Habit

Habit

Habit

Habit

Habit

Rhizome

Rhizome

Rhizome

Rhizome

Crozier

Crozier

Frond

Frond

Reduced basal pinnae

Reduced basal pinnae

Pinnae

Pinnae

Frond apex

Frond apex

Transition from full sized to reduced pinnae

Transition from full sized to reduced pinnae

Basal pinnae

Basal pinnae

Upper surface of pinna

Upper surface of pinna

Upper surface of pinna

Upper surface of pinna

Venation

Venation

Lower surface of rachis and pinnae

Lower surface of rachis and pinnae

Base of pinna and young sori

Base of pinna and young sori

Lower surface of pinna and young sori

Lower surface of pinna and young sori

Lower surface of pinna and old sori

Lower surface of pinna and old sori

Sori

Sori

Site hosted by the Royal Botanic Garden Edinburgh. Content managed by Stuart Lindsay, Gardens by the Bay, Singapore and David Middleton, Singapore Botanic Gardens. Last updated 16 November 2016.