Dryopteris cochleata (D.Don) C.Chr.

Family

Dryopteridaceae

Nomenclature

Dryopteris cochleata (D.Don) C.Chr., Index Filic.: 258. 1905; Ching, Bull. Fan Mem. Inst. Biol. 8: 434. 1938; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 312. 1941; Holttum, Dansk Bot. Ark. 20: 30. 1961; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 95. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 347, f. 31.3–31.7. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 157, 158, 202. 2000. – Nephrodium cochleatum D.Don, Prodr. Fl. Nepal.: 6. 1825. – Lastrea filix-mas var. cochleata (D.Don) Bedd., Handb. Ferns Brit. India: 250, f. 130. 1883.

Description

Rhizome short creeping; scales light brown, linear, entire, about 10 by 1 mm. Stipes stramineous, up to 30 cm in sterile and 50 cm in fertile fronds, densely scaly at base, sparsely clad with linear scales. Frond bipinnate, distinctly dimorphic; sterile lamina oblong-subdeltoid, acuminate at apex, up to 45 by 30 cm; rachis glabrescent or minutely scaly, grooved above; lateral pinnae up to 10 pairs, basal pinnae the largest or slightly smaller than the next above, slightly falcate, oblong-lanceolate, caudately acuminate at apex, narrowing towards stalked base, pinnate, up to 25 by 7 cm; pinna-rachis grooved above, winged throughout; pinnules oblong, falcate, acute to moderately acute at apex, up to 4 by 1.3 cm, shallowly lobed at margin; lobes oblique, slightly toothed; herbaceous to softly papyraceous, light green, glabrous, veins bipinnate; fertile lamina oblong or narrowly subdeltoid, up to 50 by 15 cm; pinnae ascending, linear, up to 10 by 1.5 cm; pinnules oblong, round at apex, truncate and shortly stalked or adnate at base, up to 10 by 5 mm; veins pinnate, veinlets simple or forked. Sori in one row between midrib and margin; indusia large, up to 2 mm diam., very close to each other or slightly imbricate, glabrous .

Distribution in Thailand

NORTHERN: Mae Hong Son, Chiang Mai, Chiang Rai, Lamphun, Tak; NORTH-EASTERN: Loei.

Wider Distribution

Himalayas, Burma, S China, Philippines and E Java.

Ecology

On clayey or rather dry slopes in not so dense forests at low to medium altitudes, fairly common in North, especially in the mixed forests.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Middleton et al. 4879, Chiang Mai, Doi Inthanon National Park (E); Middleton et al. 5128, Loei, Phu Suan Sai (E).

Habit

Habit

Rhizome

Rhizome

Stipe

Stipe

Fertile fronds

Fertile fronds

Crozier

Crozier

Sterile and fertile fronds

Sterile and fertile fronds

Sterile frond

Sterile frond

Fertile frond

Fertile frond

Upper surface of frond

Upper surface of frond

Young sori

Young sori

Sori

Sori

Sori

Sori

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