Arthromeris proteus (Copel.) Tagawa




Arthromeris proteus (Copel.) Tagawa, J. Jap. Bot. 27: 218. 1952; Hovenkamp, Fl. Males., Ser. II, Ferns and Fern Allies 3: 24. 1998. – Polypodium proteus Copel., Philipp. J. Sci. 1(Suppl. 2): 164. 1906. – Pleopeltis proteus (Copel.) Alderw., Bull. Dép. Agric. Indes Néerl. 27: 11. 1909. – Crypsinus proteus (Copel.) Copel., Gen. Fil.: 206. 1947. – Type: Copeland 1941 bis, Philippines, Luzon.


Description from Hovenkamp, Fl. Males., Ser. II, Ferns and Fern Allies 3: 24. 1998.
Terrestrial. Rhizome c. 4 mm thick, short creeping, phyllopodia nearly contiguous. Anatomy: vascular strands 9, without sclerified sheath, ground tissue parenchymatous, sclerified strands absent. Rhizome scales pseudopeltate, c. 1 by 3.5 mm, spreading, brown, large-celled, margin dentate. Fronds monomorphic, stipitate, pinnate. Stipes to 17 cm, lamina to 20 cm long. Lateral pinnules opposite, 1–4 pairs, 3–3.5 cm distant, fully articulate or the middle pinnae with adnate acroscopic base, 6–7 by 1–1.5 cm, base asymmetric, basiscopic side rounded, acroscopic side narrowed, terminal pinna conform, not articulate, sometimes with distinct auricles at base. Margin sinuous, cartilaginous, entire. Venation: veins distinct, connecting veins and veinlets forming areoles with free included veinlets, free veinlets simple or forked, recurrent and excurrent, recurrent in marginal areole, ending in hydathodes. Indument: sparse uniseriate, catenate, to 2 mm long hairs on the lower surface of rachis and costae. Sori on all pinnae, 1–3-seriate, in single rows between adjacent pairs of veins, round or irregularly shaped, often somewhat elongated longitudinally or spreading along the veins.

Distribution in Thailand

NORTHERN: Chiang Mai?

The exact locality is rather obscure but likely to be in Chiang Mai.

Wider Distribution

NE India, Burma, S China, Philippines.


Terrestrial on dry rocky slopes at high altitude.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threats although has only rarely been collected in Thailand.

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