Ferns of Thailand, Laos and Cambodia

Family

Athyriaceae

Nomenclature

Athyrium cuspidatum (Bedd.) M.Kato, Bot. Mag. (Tokyo) 90: 27. 1977. – Lastrea cuspidata Bedd., Ferns Brit. India: t. 118. 1870; Bedd., Handb. Ferns Brit. India: 232, f. 119. 1883, based on Aspidium cuspidatum Mett., Abh. Senckenberg. Naturf. Ges. 2: 376. 1858, non Desv., 1827. – Microchlaena cuspidata (Bedd.) Ching, Acta Phytotax. Sin. 9: 99. 1964; Tagawa & K.Iwats., Acta Phytotax. Geobot. 22: 103. 1967; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 101. 1967; K.Iwats., Acta Phytotax. Geobot. 24: 188. 1970. – Kuniwatsukia cuspidata (Bedd.) Pic.-Serm., Webbia 28: 455. 1973; Tagawa & K.Iwats., Fl. Thailand 3: 442, f. 47.1–4. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 198. 2000. – Anisocampium cuspidatum (Bedd.) Yea C.Liu, W.L.Chiou & M.Kato, Taxon 60(3): 829. 2011.

Aspidium yunnanense Christ, Bull. Herb. Boiss. 6: 965. 1898. – Microchlaena yunnanensis (Christ) Ching, Bull. Fan Mem. Inst. Biol. 8: 325. 1938; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 87. 1965.

Description

Terrestrial. Rhizome short, creeping to ascending; scales linear-lanceolate, gradually narrowing towards the tailed apex, up to 12 by 1.2 mm, entire. Stipes purplish when young, stramineous to pale brown, up to 60 cm long, broadened near scaly base. Laminae imparipinnate, oblong in outline, about 60 by 35 cm; pinnae linear, acuminate at apex, truncate to auricled at acroscopic and cuneate at basiscopic bases, shallowly lobed at margin, up to 20 by 1.3–1.7 cm, the soriferous pinnae usually narrower; lobes triangular, with sharp teeth at apex; papyraceous, green, paler beneath; rachis and costa glabrous or with linear scales; veins pinnate, the veinlets simple, all except the basal pairs reaching the margin of lobes. Sori dorsal on veinlets, costular, with small round-reniform indusia lacerate at margin .

Distribution in Thailand

NORTHERN: Chiang Mai, Chiang Rai, Lamphun.

Wider Distribution

Nepal, Assain, Upper Burma, and SW China.

Ecology

On rather dry mountain slopes in mixed or evergreen forests at 800–1600 m alt.

Similar species

Dryopteris hirtipes (Blume) Kuntze

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threats.

Voucher specimens - Thailand

Middleton et al. 5006, Chiang Mai, Doi Chiang Dao Wildlife Sanctuary (E).

	Ferns of Thailand, Laos and Cambodia
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Athyrium cuspidatum (Bedd.) M.Kato Family Athyriaceae Nomenclature Athyrium cuspidatum (Bedd.) M.Kato, Bot. Mag. (Tokyo) 90: 27. 1977. – Lastrea cuspidata Bedd., Ferns Brit. India: t. 118. 1870; Bedd., Handb. Ferns Brit. India: 232, f. 119. 1883, based on Aspidium cuspidatum Mett., Abh. Senckenberg. Naturf. Ges. 2: 376. 1858, non Desv., 1827. – Microchlaena cuspidata (Bedd.) Ching, Acta Phytotax. Sin. 9: 99. 1964; Tagawa & K.Iwats., Acta Phytotax. Geobot. 22: 103. 1967; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 101. 1967; K.Iwats., Acta Phytotax. Geobot. 24: 188. 1970. – Kuniwatsukia cuspidata (Bedd.) Pic.-Serm., Webbia 28: 455. 1973; Tagawa & K.Iwats., Fl. Thailand 3: 442, f. 47.1–4. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 198. 2000. – Anisocampium cuspidatum (Bedd.) Yea C.Liu, W.L.Chiou & M.Kato, Taxon 60(3): 829. 2011. Aspidium yunnanense Christ, Bull. Herb. Boiss. 6: 965. 1898. – Microchlaena yunnanensis (Christ) Ching, Bull. Fan Mem. Inst. Biol. 8: 325. 1938; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 87. 1965. Description Terrestrial. Rhizome short, creeping to ascending; scales linear-lanceolate, gradually narrowing towards the tailed apex, up to 12 by 1.2 mm, entire. Stipes purplish when young, stramineous to pale brown, up to 60 cm long, broadened near scaly base. Laminae imparipinnate, oblong in outline, about 60 by 35 cm; pinnae linear, acuminate at apex, truncate to auricled at acroscopic and cuneate at basiscopic bases, shallowly lobed at margin, up to 20 by 1.3–1.7 cm, the soriferous pinnae usually narrower; lobes triangular, with sharp teeth at apex; papyraceous, green, paler beneath; rachis and costa glabrous or with linear scales; veins pinnate, the veinlets simple, all except the basal pairs reaching the margin of lobes. Sori dorsal on veinlets, costular, with small round-reniform indusia lacerate at margin . Distribution in Thailand NORTHERN: Chiang Mai, Chiang Rai, Lamphun. Wider Distribution Nepal, Assain, Upper Burma, and SW China. Ecology On rather dry mountain slopes in mixed or evergreen forests at 800–1600 m alt. Similar species Dryopteris hirtipes (Blume) Kuntze Proposed IUCN Conservation Assessment Least Concern (LC). This species is widespread and not under any known threats. Voucher specimens - Thailand Middleton et al. 5006, Chiang Mai, Doi Chiang Dao Wildlife Sanctuary (E). Rhizome Scales on rhizome Upper surface of lamina Upper surface of base of terminal pinna Upper surface of pinna Lower surface of lamina Lower surface of pinna and sori
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Athyrium cuspidatum (Bedd.) M.Kato