Ferns of Thailand, Laos and Cambodia > Cyclosorus articulatus

Family

Thelypteridaceae

Nomenclature

Cyclosorus articulatus (Houlston & T.Moore) Panigrahi, Res. J. Pl. Environ. 9: 66. 1993. – Nephrodium articulatum Houlston & T.Moore, Gard. Mag. Bot. 1851: 293, based on Aspidium articulatum Lowe, Ferns Br. Exot. 6, t. 49. 1857, non Sw. 1801. – Pronephrium articulatum (Houlston & T.Moore) Holttum, Blumea 20: 116. 1972. – Thelypteris articulata (Houlston & T.Moore) Tagawa & K.Iwats., Acta Phytotax. Geobot. 26: 169. 1975; Tagawa & K.Iwats., Fl. Thailand 3: 406, f. 41.3 & 42.6–9. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 225. 2000.

Nephrodium glandulosun var. laete-strigosum C.B.Clarke, Trans. Linn. Soc. London, Bot. 1: 532, t. 74, f. 2. 1880. – Dryopteris indica Alderw., Malayan Ferns: 224. 1909. – Cyclosorus laete-strigosus (C.B.Clarke) Ching, Bull. Fan Mem. Inst. Biol. 8: 227. 1938, nom. superfl. – Thelypteris indica (Alderw.) Reed, Phytologia 17: 284. 1968; Tagawa & K.Iwats., Acta Phytotax. Geobot. 25: 17. 1971.

Description

Rhizome short, massive, short creeping to ascending; scales hairy. Stipes up to 87 cm long, hairy. Laminae indistinctly dimorphic with 15–26 pairs of pinnae; lamina oblong-lanceolate, up to 150 by 60cm; sterile pinnae oblong-lanceolate, gradually narrowing towards long-acuminate apex, narrowly cuneate at basiscopic and subtruncate at acroscopic bases, shallowly lobed, up to 30.5 by 4 cm; soriferous pinnae narrower, cuneate or broadly cuneate at base, very shortly stalked or sessile, up to 20 by 2.5 cm, shallowly lobed; lobes semi-orbicular, or obtuse at apex; papyraceous to softly chartaceous, glabrous, with spherical sessile glands or not; veins meniscioid with a few pairs of free veinlets near margin. Sori medial, round reniform to elongate reniform; indusia large, thin but firm, persistent, glabrous .

Distribution in Thailand

NORTHERN: Lampang, Tak; SOUTH-WESTERN: Kanchanaburi.

Wider Distribution

Sri Lanka, Himalaya, Burma, and W China.

Ecology

Moist evergreen forests along streamlets at medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This variety is widespread and not under any known threats.

Voucher specimens - Thailand

Middleton et al. 4764, Kanchanaburi, Khao Laem National Park (E).

Habit

Rhizome

Growing tip of rhizome

Crozier

Frond apex

Venation

Old sori

Young sori

	Ferns of Thailand, Laos and Cambodia
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Cyclosorus articulatus (Houlston & T.Moore) Panigrahi Family Thelypteridaceae Nomenclature Cyclosorus articulatus (Houlston & T.Moore) Panigrahi, Res. J. Pl. Environ. 9: 66. 1993. – Nephrodium articulatum Houlston & T.Moore, Gard. Mag. Bot. 1851: 293, based on Aspidium articulatum Lowe, Ferns Br. Exot. 6, t. 49. 1857, non Sw. 1801. – Pronephrium articulatum (Houlston & T.Moore) Holttum, Blumea 20: 116. 1972. – Thelypteris articulata (Houlston & T.Moore) Tagawa & K.Iwats., Acta Phytotax. Geobot. 26: 169. 1975; Tagawa & K.Iwats., Fl. Thailand 3: 406, f. 41.3 & 42.6–9. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 225. 2000. Nephrodium glandulosun var. laete-strigosum C.B.Clarke, Trans. Linn. Soc. London, Bot. 1: 532, t. 74, f. 2. 1880. – Dryopteris indica Alderw., Malayan Ferns: 224. 1909. – Cyclosorus laete-strigosus (C.B.Clarke) Ching, Bull. Fan Mem. Inst. Biol. 8: 227. 1938, nom. superfl. – Thelypteris indica (Alderw.) Reed, Phytologia 17: 284. 1968; Tagawa & K.Iwats., Acta Phytotax. Geobot. 25: 17. 1971. Description Rhizome short, massive, short creeping to ascending; scales hairy. Stipes up to 87 cm long, hairy. Laminae indistinctly dimorphic with 15–26 pairs of pinnae; lamina oblong-lanceolate, up to 150 by 60cm; sterile pinnae oblong-lanceolate, gradually narrowing towards long-acuminate apex, narrowly cuneate at basiscopic and subtruncate at acroscopic bases, shallowly lobed, up to 30.5 by 4 cm; soriferous pinnae narrower, cuneate or broadly cuneate at base, very shortly stalked or sessile, up to 20 by 2.5 cm, shallowly lobed; lobes semi-orbicular, or obtuse at apex; papyraceous to softly chartaceous, glabrous, with spherical sessile glands or not; veins meniscioid with a few pairs of free veinlets near margin. Sori medial, round reniform to elongate reniform; indusia large, thin but firm, persistent, glabrous . Distribution in Thailand NORTHERN: Lampang, Tak; SOUTH-WESTERN: Kanchanaburi. Wider Distribution Sri Lanka, Himalaya, Burma, and W China. Ecology Moist evergreen forests along streamlets at medium altitudes. Proposed IUCN Conservation Assessment Least Concern (LC). This variety is widespread and not under any known threats. Voucher specimens - Thailand Middleton et al. 4764, Kanchanaburi, Khao Laem National Park (E). Habit Rhizome Growing tip of rhizome Crozier Frond apex Venation Old sori Young sori Young sori
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Cyclosorus articulatus (Houlston & T.Moore) Panigrahi