Family

Thelypteridaceae

Nomenclature

Cyclosorus asperus (C.Presl) B.K.Nayar & S.Kaur, Companion Beddome's Handb. Ferns Brit. India 66. 1974. – Goniopteris aspera C.Presl, Tent. Pterid.: 183. 1836, based on Polypodium asperum C.Presl, Rel. Haenk. 1: 24, t. 3, f. 4.,1825, non L. 1753. – Thelypteris aspera (C.Presl) K.Iwats., Mem. Coll. Sci. Univ. Kyoto B. 31: 192. 1965; Tagawa & K.Iwats., Acta Phytotax. Geobot. 25: 17. 1971; Tagawa & K.Iwats., Fl. Thailand 3: 411. 1988. – Pronephrium asperum (C.Presl) Holttum, Blumea 20: 112. 1972; Holttum, Fl. Males., Ser. II, Pterid. 1: 512, f. 14g–i. 1982 [‘1981’]; Boonkerd & Pollawatn, Pterid. Thailand: 225. 2000. – Type: Haenke, Philippines, Luzon (PRC).

Dryopteris gymnopteridifrons Hayata, Ic. Pl. Formos. 8: 148, f. 75, 76. 1919. – Pronephrium gymnopteridifrons (Hayata) Holttum, Blumea 20: 112 1972.

Abacopteris multilineata var. malayensis Holttum, Rev. Fl. Malaya ed. 1, 2: 297. 1955 [‘1954’], nom. illeg. – Thelypteris multilineata var. malayensis Reed, Phytologia 17: 294. 1968.

Thelypteris urophylla auct. non (Mett.) K.Iwats.: Tagawa & K.Iwats., SouthE Asian Stud. 5: 71. 1967, p.p.

Description

Rhizome creeping; scales oblong-subtriangular, about 7 by 1.2 mm, pale brown, hairy at margin. Stipes 35–90 cm long, dark and scaly at base, stramineous or slightly flushed red. Laminae oblong, 30–80 by 20–44 cm; lateral pinnae about 4–6(–8) pairs, lowest pair sometimes reduced, upper ones larger, sessile, narrowly oblong, caudate at apex, gradually narrowing towards narrowly cuneate base, 15–37 by 2.5–6.3 cm, subentire or irregularly undulate, cartilaginous at margin; venation meniscioid; texture papyraceous, firm, pale green to green, verrucose, sparsely pubescent or glabrescent. Sori in two rows, medial one close to excurrent veinlets; indusia small, persistent but usually immersed among sporangia at maturity, hairy; sporangia with a few setose hairs.

Distribution in Thailand

NORTHERN: Lampang; NORTH-EASTERN: Loei; SOUTH-EASTERN: Chanthaburi; PENINSULAR: Yala.

Wider Distribution

S China, Taiwan, throughout Malesia to N Queensland.

Ecology

Terrestrial on rather dry slopes in evergreen forests along streamlets at medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Notes

Although the material from Thailand has setose hairs on the sporangia Holttum (1982) notes that the type from the Philippines lacks setose hairs on the sporangia .

Voucher specimens - Thailand

Middleton et al. 4693, Chanthaburi, Khlong Khrue Wai Wildlife Sanctuary (E).