Drynaria quercifolia (L.) J.Sm.




Drynaria quercifolia (L.) J.Sm., J. Bot. 3: 398. 1841; Bedd., Handb. Ferns Brit. India: 341, f. 191. 1883; E.Smith, J. Siam Soc. Nat. Hist. Suppl. 8: 6. 1929; C.Chr., Contr. U.S. Natl. Herb. 26: 335. 1931; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 518. 1941; Holttum, Rev. Fl. Malaya ed. 1, 2: 182, f. 88. 1955 [‘1954’]; Holttum, Dansk Bot. Ark. 23: 231. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 77. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 59. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 546. 1989; Hovenkamp & Roos, Fl. Males., Ser. II, Ferns and Fern Allies 3: 40. 1998; Boonkerd & Pollawatn, Pterid. Thailand: 269. 2000; Newman et al., Checkl. Vasc. Pl. Lao PDR: 29. 2007; Leti et al., Fl. Photogr. Cambodge 557. 2013. – Polypodium quercifolium L., Sp. Pl. 2: 1087. 1753. – Type: Herb. Hermann I: 39 (BM).


Rhizome short creeping, 1–2 cm diam., densely scaly; scales dark brown, gradually narrowing from base to tailed apex, up to 4–18 mm long, 1.3 mm broad, the margin paler and densely toothed. Nest-leaves sessile, ovate, 15–32(–50) by 10–20(–40) cm, shallowly lobed; lobes close, 5 by 3 cm, rounded to moderately acute at apex, entire. Foliage-leaves: stipes 15–25(–35) cm long, stramineous, very narrowly winged throughout, densely scaly at base, these similar to the rhizome scales; laminae pinnatifid, oblong to narrower, 40–80(–150) by 15–50 cm, deeply pinnatifid, continuous to the next ones by rather broad wings more than 1 cm in breadth; lobes ascending, gradually narrowing from base to acute or acuminate apex, entire and more or less crisped at margin, 1–25(–30) by 2–4.5 cm; veins distinct on both surfaces, finely anastomosing with more than 10 rows of areoles between main veins; coriaceous, shining pale green, glabrous. Sori round or oblong, two rows between adjacent main veins.

Distribution in Thailand

NORTHERN: Chiang Mai, Chiang Rai, Tak, Phitsanulok; NORTH-EASTERN: Nong Khai; SOUTH-WESTERN: Kanchanaburi, Prachuap Khiri Khan; SOUTH EASTERN: Chon Buri, Chanthaburi, Trat; PENINSULAR: Phangnga, Surat Thani, Krabi, Nakhon Si Thammarat, Phuket, Phatthalung, Trang, Satun, Yala.

Distribution in Laos

Champasak, Xieng Khouang.

Distribution in Cambodia

Mondulkiri, Siem Reap.

Wider Distribution

Sri Lanka, India to S China and Indochina, throughout Malesia to Fiji and tropical Australia.


On rather dry rocks on hillsides in light shade or at edge of forests, fairly common at low altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Middleton et al. 4450, Krabi, Ko Phi Phi National Park (E).

Voucher specimens - Laos

Maxwell 97-1134, Champasak (CMU)

Voucher specimens - Cambodia

Long et al. CL342, Mondulkiri (P).





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