Lindsaea javanensis Blume




Lindsaea javanensis Blume, Enum. Pl. Javae: 219. 1828; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 124. 1939; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 75. 1967; Kramer, Fl. Males., Ser. II, Pterid. 1: 208. 1971; Kramer, Gard. Bull. Singapore 26: 25. 1972; Tagawa & K.Iwats., Fl. Thailand 3: 134. 1985; Boonkerd & Pollawatn, Pterid. Thailand: 92. 2000. – Schizoloma javanense (Blume) Holttum, Rev. Fl. Malaya ed. 1, 2: 349, f. 202. 1955 [‘1954’].

Lindsaea flabellata var. gigantea Hook., Sp. Fil. 1: 211, t. 63C. 1864. – Lindsaea tenera var. gigantea (Hook.) Holttum, Gard. Bull. Straits Settlem. 5: 65. 1930.

Lindsaea commixta auct. non Tagawa: Newman et al., Checkl. Vasc. Pl. Lao PDR: 27. 2007.


Description from Kramer, Fl. Males., Ser. II, Pterid. 1: 208. 1971.
Rhizome short creeping, 1–2 mm diam.; scales medium brown, lanceolate, long-acuminate, to 1.5 mm long, to 4-seriate at base, with a long uniseriate apex. Fronds clustered; stipes 10–30 cm long, especially in large leaves much longer than the lamina, dark reddish brown to atropurpureous, ± lustrous, abaxially terete at base, upward gradually obtusely, at apex mostly acutely bi-angular, the angles and especially the borders of the adaxial groove often pale. Laminae herbaceous to chartaceous, dark green or olivaceous when dry, very variable, triangular or oblong or in small leaves sometimes transversely triangular, 7–20 cm long, 6–15 cm wide, 1.25 to (in large leaves) 1.75 times as long as wide, simply pinnate to amply bipinnate, in the first case with up to as little as 2 pairs of lateral pinnules and a distinct but not conform terminal one, in the second case with up to 5 (pinnate) pinnae to a side and several simple ones above, which are gradually reduced to and confluent (or not) with the terminal pinnule (segment). Primary rachis dark, adaxially with a narrow, pale-edged groove, abaxially flattened, pale-angled. Primary pinnae their width apart to contiguous, spreading or in plurijugate bipinnate leaves ascending. Pinnules of simply pinnate or subbipinnate leaves rhombic, very often with long-acuminate to caudate apex, with very unequal base, basiscopically much more cut away, to c. 8 by 2 cm; apex of lamina similar but ± symmetric at base; apices of fully pinnate pinnae similar but smaller, progressively smaller and less pointed as there are more secondary pinnules; larger transitional pinnules between pinnate pinnae and the lamina apex often rhombic-caudate, smaller ones subtrapeziform or subflabellate, very obtuse; pinnules of fully pinnate pinnae up to 8 to a side, mostly not contiguous, subtrapeziform, rounded-rhombic or subflabellate. Secondary rachises abaxially flattened or slightly sulcate, abruptly pale at their insertion on the dark primary rachis, usually distinctly green-margined. Fertile pinnule-margin subentire or minutely erose, in larger leaf segments here and there incised by shallow crenations; sterile margin, especially near the segment bases, sharply serrate or dentate, with deeper incisions. Lobes of pointed pinnules often slightly concave. Veins immersed but ± evident, free, mostly twice forked, c. 1 mm apart; larger pointed pinnules and terminal divisions with a percurrent costa. Sori continuous in small pinnules, progressively more interrupted in larger ones, bi- to plurinerval. Indusium pale, subentire to erose, 0.3–0.4 mm wide, almost reaching the margin to falling short of it by more than its width, little reflexed at maturity. Spores light brown, trilete , smooth, c. 25?. {EXPLAIN?}

Distribution in Thailand

NORTH-EASTERN: Loei; CENTRAL: Nakhon Nayok; PENINSULAR: Nakhon Si Thammarat, Trang, Yala.

Distribution in Laos


Distribution in Cambodia

Kompong Chhnang

Wider Distribution

India (Assam), Burma, Indochina and W Malesia, north to the Ryukyus and southern edge of Japan.


Terrestrial on mountain slopes at medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Middleton et al. 3678A, Yala, Hala-Bala Wildlife Sanctuary (E).

Voucher specimens - Laos

Newman et al. 867, Khammouane, Nam Theun (E, P).

Voucher specimens - Cambodia

Poilane 28795, Kompong Chhnang (P).

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