Microsorum zippelii (Blume) Ching




Microsorum zippelii (Blume) Ching, Bull. Fan Mem. Inst. Biol. 4: 308. 1933; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 479. 1941; Holttum, Rev. Fl. Malaya ed. 1, 2: 176, f. 85. 1955 [‘1954’]; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 52. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 525. 1989; Nooteboom, Blumea 42: 371. 1997; Bosman et al., Fl. Males., Ser. II, Ferns and Fern Allies 3: 130. 1998; Boonkerd & Pollawatn, Pterid. Thailand: 281. 2000. – Polypodium zippelii Blume, Fl. Javae Fil.: 172, t. 80. 1829. – Pleopeltis zippelii (Blume) T.Moore, Index Fil.: 348. 1862; Bedd., Handb. Ferns Brit. India: 357. 1883. – Neocheiropteris zippelii (Blume) Bosman, Leiden Bot. Ser. 14: 123. 1991. – Neolepisorus zippelii (Blume) Li Wang, Bot. J. Linn. Soc. 162(1): 36. 2010. – Type: Zippelius s.n. (L), Java.


Rhizome creeping, 1–5 mm diam., bearing fronds more than 1 cm apart, densely scaly throughout; scales pseudopeltate, narrowly oblong-subtriangular, gradually narrowing towards attenuate apex, 2.5–7 by 1–2 mm, dark brown, clathrate, toothed at margin. Stipes (0.8–)5–8 cm long, indistinct on the upper, winged portion, scaly at base, green to stramineous. Laminae oblong-lanceolate, caudately acuminate at apex, broadest at middle portion, gradually narrowing towards long-attenuate base, entire and flat at margin, 6.5–65 by 1–8 cm; midrib and main lateral veins raised on both surfaces, veins more or less distinct, anastomosing, smaller areoles in 3 rows between adjacent main veins with branched included veinlets; papyraceous. Sori round , usually at junction of veinlets, arranged in two rows between main veins, about 2 mm diam.

Distribution in Thailand

NORTHERN: Chiang Mai, Chiang Rai; NORTH-EASTERN: Loei; SOUTH-WESTERN: Phetchaburi.

Distribution in Laos


Distribution in Cambodia

[Pnom Chom]

Wider Distribution

Himalayas and throughout Malesia, northeast to S China (Hainan) and Indochina.


In muddy crevices of rocks or on mountain slopes by streams in dense evergreen forests at high altitudes, rather rare.

Similar species

Leptochilus hemionitideus (C.Presl) Noot.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.


The sori are placed between the smaller areoles, thus arranging in two rows between the main lateral veins. In this species, therefore, the regular arrangement of sori in two rows is based upon the pattern of venation and thus a distinct specific feature. The size of Thai plants is smaller in comparison with the plants from the other regions.

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