Pleocnemia submembranacea (Hayata) Tagawa & K.Iwats.




Pleocnemia submembranacea (Hayata) Tagawa & K.Iwats., Acta Phytotax. Geobot. 26: 61. 1974; Tagawa & K.Iwats., Fl. Thailand 3: 386. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 207. 2000. – Aspidium submembranaceum Hayata, Ic. Pl. Formos. 4: 188, f. 126. 1914. – Type: Hayata & Sasaki s.n. (TI), Urai, Taiwan.

Pleocnemia winitii Holttum, Reinwardtia 1: 181. 1951; Holttum, Kew Bull. 29: 354. 1974; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 100. 1967. – Type: Winit 36 (SING), Lampang.

Tectaria leuzeana auct. non (Gaudich.) Copel.: Bedd., Handb. Ferns Brit. India: 228. 1883, p.p.; C.Chr., Contr. U.S. Natl. Herb. 26: 331. 1931; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 405. 1941.


Rhizome thick, short, erect, scaly near apex; scales about 20 by 1 mm, dark brown, more or less crisped, entire or irregularly toothed at margin. Stipes castaneous and densely scaly at base, stramineous, pubescent or glabrescent upwards. Laminae more than 1 m long, tripinnatifid; pinnae asymmetrically oblong-subdeltoid to oblong-lanceolate, acuminate at apex; rachis and pinna-rachis pubescent on upper surface; basal basiscopic pinnules of lower pinnae sometimes enlarged, pinnate with deeply lobed secondary pinnules; the other pinnules subtriangular, widest at base, gradually narrowing towards caudate apex, commonly about 10 by 3.5 cm, stalked, lobed to 4/5 way towards costule, the lower segments sometimes free; segments oblong-subdeltoid, moderately acute at apex, more or less serrate at margin; chartaceous, green, brownish in dried specimens; veins anastomosing forming costal and costular areoles, or with one more row of areoles outside the costular ones, sparsely hairy beneath. Sori dorsal on veins, in one to two rather irregular rows at each side of midrib, exindusiate, sporangia with glands.

Distribution in Thailand

NORTHERN: Chiang Mai, Chiang Rai, Lampang, Phitsanulok.

Distribution in Laos


Wider Distribution

N India, Burma, Indochina, China (Hainan, Hong Kong) and Taiwan.


On humus rich mountain slopes usually near streams in dense forests at medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

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