Cyclosorus heterocarpus (Blume) Ching

Family

Thelypteridaceae

Nomenclature

Cyclosorus heterocarpus (Blume) Ching, Bull. Fan. Mem. Inst. Biol. 8: 180. 1938; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 385. 1941; Holttum, Rev. Fl. Malaya ed. 1, 2: 269, f. 155. 1955 [‘1954’]. – Aspidium heterocarpum Blume, Enum. Pl. Javae.: 155. 1828. – Nephrodium heterocarpum (Blume) T.Moore, Ind. Fil. 93. 1958. – Dryopteris heterocarpa (Blume) Kuntze, Rev. Gen. Pl. 2: 813. 1891. – Thelypteris heterocarpa (Blume) Morton, Amer. Fern J. 49: 113. 1959; Tagawa & K.Iwats., Fl. Thailand 3: 416. 1988; Newman et al., Checkl. Vasc. Pl. Lao PDR: 33. 2007. – Sphaerostephanos heterocarpus (Blume) Holttum in B.K.Nayar & S.Kaur, Companion Handb. Ferns Brit. India: 209. 1974; Holttum, Kalikasan 4: 67. 1975; Holttum, Fl. Males., Ser. II, Pterid. 1: 457, f. 1q. 1982 [‘1981’]; Boonkerd & Pollawatn, Pterid. Thailand: 229. 2000. – Type: Blume s.n., Java, ad pedem Boerangrang (L, sheet 908,332-1158).

Nephrodium glaucostipes Bedd., Suppl. Ferns Brit. Ind.: 80. 1892.

Description

Rhizome erect; scales narrow, up to 8 by 1.2 mm, hirsute, dark brown. Stipes 24–50 cm long, bearing reduced pinnae nearly to base, hirsute. Laminae 42–57.5 x 14–24.5 cm, oblong-lanceolate, acute at apex, 16–30 pairs of free full-sized primary pinnae, lower pinnae suddenly reduced in size to mere auricles, full-sized pinnae nearly straight, ascending, linear-lanceo1ate, caudate-acuminate at apex, cuneate at sessile base, 8–20 by 1.1–1.8(–2.5) cm, 2–4 cm apart at base, lobed to half-way towards costa; segments oblong, oblique, rounded to obtuse at apex; thinly papyraceous, green to yellowish-green, pubescent on upper surface, glandular below; rachis and costa hairy throughout; veins pinnate, veinlets simple, 1.5 basal pairs uniting below sinus. Sori medial; indusia persistent, brown at maturity, glabrous or sparsely hairy .

Distribution in Thailand

SOUTH-EASTERN: Chanthaburi; PENINSULAR: Surat Thani, Phangnga, Nakhon Si Thammarat, Trang, Satun, Yala, Narathiwat.

Distribution in Cambodia

Kampot

Wider Distribution

S China, Vietnam, throughout Malesia to Polynesia.

Ecology

Terrestrial in evergreen forests at low altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Middleton et al. 5451, Phangnga, Sra Nang Manora Forest Park (E); Shimizu et al. T-8186, Nakhon Si Thammarat, Thung Song, Khao Kao (K).

Voucher specimens - Laos

Unspecified

Habit

Habit

Rhizome

Rhizome

Crozier

Crozier

Reduced basal pinnae

Reduced basal pinnae

Pinnae

Pinnae

Frond apex

Frond apex

Venation

Venation

Position of sori on veins

Position of sori on veins

Lower surface of pinna

Lower surface of pinna

Lower surface of rachis and pinna

Lower surface of rachis and pinna

Lower surface of rachis and pinnae

Lower surface of rachis and pinnae

Sori

Sori

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