Cyclosorus tylodes (Kunze) Panigrahi

Family

Thelypteridaceae

Nomenclature

Cyclosorus tylodes (Kunze) Panigrahi, Res. J. Pl. Environ. 9: 67. 1993. – Aspidium tylodes Kunze, Linnaea 24: 283. 1851. – Lastrea ochthodes var. tylodes (Kunze) Bedd., Handb. Ferns Brit. India: 240. 1883. – Thelypteris tylodes (Kunze) Ching, Bull. Fan Mem. Inst. Biol. 6: 296. 1936; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 357. 1941; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 67. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 421. 1988, as Thelypteris xylodes; Boonkerd & Pollawatn, Pterid. Thailand: 231. 2000, as Thelypteris xylodes. – Pseudocyclosorus tylodes (Kunze) Ching, Acta Phytotax. Sin. 8: 324. 1963; Holttum, Fl. Males., Ser. II, Pterid. 1: 413, f. 10d–e. 1982 [‘1981’]. – Syntypes: Schmid-Koch 11, 63, 89, 115, Nilgiri Hills, S. India.

Dryopteris tuberculifera C.Chr., Contr. U.S. Natl. Herb. 26: 275. 1931. – Thelypteris tuberculifera (C.Chr.) Ching, Bull. Fan Mem. Inst. Biol. 6: 295. 1936; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 78. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 67. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 421. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 231. 2000. – Cyclosorus tuberculifer (C.Chr.) Panigrahi, Res. J. Pl. Environ. 9: 67. 1993.

Description

Rhizome short, ascending to suberect; scales small, caducous, up to 3 by 0.8 mm, brown, hairy. Stipes 40–80 cm long, variable in size, bearing auricles of reduced pinnae nearly to the base. Laminae oblong-lanceolate, acute to acuminate at apex, 15–110 by 15–55 cm, variable in size; pinnae 16–31 pairs, 7.8–30 x 1.2–2.5 cm, lower pinnae suddenly reduced to auricles, lowest normal pinna often reduced and deflexed, middle pinnae linear-lanceolate, gradually narrowing towards long-acuminate apex, broadly cuneate at sessile base, deeply lobed to 3/4 way to costa; segments narrowly oblong, oblique, acute at apex, entire or crenate; chartaceous, deep green; veins pinnate, veinlets simple, basal ones running to callous-membrane in sinus between segments; costa hairy above and glabrous beneath, lamina and veinlets glabrous above and beneath, small hairs around margin. Sori round, medial to costular; indusia persistent, glabrous .

Distribution in Thailand

NORTHERN: Chiang Mai, Chiang Rai, Tak, Phitsanulok; NORTH-EASTERN: Loei.

Wider Distribution

Sri Lanka, S India, Himalaya, S & SW China and Vietnam.

Ecology

On wet sandy ground or stream-beds in light shade at medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Notes

We are following the species concept of Holttum, Kew Bulletin 34: 499–516(1979) rather than that of the Flora of Thailand account in which Thelypteris tuberculifera is recognised as a separate species .

Voucher specimens - Thailand

Middleton et al. 5126, Loei, Phu Suan Sai National Park (E).

Upper surrface of pinnae

Upper surrface of pinnae

Lower surface of pinnae

Lower surface of pinnae

Upper surface of rachis and pinnae

Upper surface of rachis and pinnae

Lower surface of rachis and pinnae

Lower surface of rachis and pinnae

Lower surface of lamina with mature sori

Lower surface of lamina with mature sori

Lower surface of pinnae and young sori

Lower surface of pinnae and young sori

Mature sori

Mature sori

Sori

Sori

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