Coniogramme fraxinea (D.Don) Diels


Pteridaceae, subfamily Cryptogrammoideae


Coniogramme fraxinea (D.Don) Diels, Nat. Pflanzenfam. 1(4): 262. 1899; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 191. 1940; Holttum, Rev. Fl. Malaya ed. 1, 2: 589. 1955 [‘1954’]; Tagawa & K.Iwats., Fl. Thailand 3: 190. 1985. – Diplazium fraxineum D.Don, Prodr. Fl. Nepal.: 12. 1825; Newman et al., Checkl. Vasc. Pl. Lao PDR: 31. 2007. – Syngramma fraxinea (D.Don) Bedd., Handb. Ferns Brit. India: 386, f. 222. 1883.

Coniogramme fraxinea (D.Don) Diels var. serrulata (Blume) Hieron., Hedwigia 57: 289. 1916; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 191. 1940; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 107. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 190. 1985; Boonkerd & Pollawatn, Pterid. Thailand: 100, 122. 2000; Newman et al., Checkl. Vasc. Pl. Lao PDR: 31. 2007. – Gymnogramma serrulatum Blume, Enum. Pl. Javae.: 113. 1828; Newman et al., Checkl. Vasc. Pl. Lao PDR: 31. 2007.


Rhizome creeping, thick, densely covered with scales. Stipes stramineous, up to 1 m or more long, 1 cm diam., usually smaller, densely scaly at base, sparsely clad with minute scales upwards. Laminae bipinnate, up to 80 by 60 cm; rachis and pinna-rachis like the upper part of stipe, stramineous, minutely scaly, grooved on upper surface, grooves decurrent; lateral pinnae 5–8 pairs, lower 3–5 pairs bipinnate, upper ones simple; pinnules of larger pinnae like the upper simple lateral pinnae and apical ones, the lowest anterior pinnules rarely trifoliate or pinnate; leaflets oblong or narrower or oblong-lanceolate, usually slightly falcate, caudate at apex, cuneate at stalked base, dentate at margin, 20–30 by 3–4.5 cm; the apical pinnae or pinnules sometimes larger, herbaceous, thin, glabrous; costa distinctly raised on lower surface, grooved on upper surface; veins a few times forked, hydathodes placed below the bottom of sinus. Sori extending along veins, exindusiate .

Distribution in Thailand

NORTHERN: Chiang Mai, Lampang, Tak; NORTH-EASTERN: Loei; SOUTH-EASTERN: Chanthaburi.

Distribution in Laos

Hua Phan.

Wider Distribution

Laos, Sumatra and Java. The range of the species is in Sri Lanka, India, S China to Taiwan, Indochina and Malesia to the Philippines.


On rather moist mountain slopes in dense evergreen forests at high altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any immediate known threat.


Coniogramme fraxinea var. serrulata was previously distinguished from the type variety in having the margins of the lobes serrulate.  We find this character to be continuous .

Voucher specimens - Thailand

Middleton et al. 4943, Chiang Mai, Doi Pha Ti Do (E).

Voucher specimens - Laos

Poilane 2121, Hua Phan, Sam Nuea (P).





Lower surface of lamina

Lower surface of lamina



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