Davallia divaricata Blume




Davallia divaricata Blume, Enum. Pl. Javae: 237. 1828; Bedd., Handb. Ferns Brit. India: 60. 1883; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 107. 1939; Holttum, Rev. Fl. Malaya ed. 1, 2: 362, f. 209. 1955 [‘1954’]; Holttum, Dansk Bot. Ark. 23: 234. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 76. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 161. 1985; Nooteboom, Fl. Males., Ser. II, Ferns and Fern Allies 3: 252. 1998; Boonkerd & Pollawatn, Pterid. Thailand: 233. 2000. – Araiostegia divaricata (Blume) Kato, Acta Phytotax. Geobot. 26: 158. 1975. – Wibelia divaricata (Blume) M.Kato & Tsutsumi, Acta Phytotax. Geobot. 59(1): 13. 2008. – Type: Blume s.n. (L sheet 908.333-153 holo; K), Java, G. Burangan.

Davallia orientalis C.Chr., Bull. Dept. Biol. Sun Yatsen Univ. 3: 104, t. 43. 1932. – Davallia divaricata var. orientalis (C.Chr.)  Tardieu & C.Chr., Notul. Syst. (Paris) 6: 3, pl.1, 3-4. 1937; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 107. 1939.


Rhizome creeping, thick, usually more than 8 mm diam., densely scaly throughout; scales linear-lanceolate, gradually narrowing towards long-tailed apex, up to 15 by 2 mm, entire, minutely toothed, brown, orange-brown or pale brown. Stipes castaneous to darker, up to 50(–60) cm long, terete, minutely scaly, Laminae oblong, acuminate at apex, quadripinnate to quadripinnatifid, less divided in sterile fronds, up to 80(–100) by 65(70) cm; all pinnae anadromous; basal pinnae the largest, narrowly deltoid, broadly cuneate at base, stalked, up to 40 by 30 cm; pinnules gradually narrowing from base to long caudate-acuminate apex, unequally cuneate at base, stalked; ultimate segments round to moderately acute, with acute entire lobes, subcoriaceous, glabrous; veins pinnate, without false veinlets. Sori small, terminal on veinlets, the edges 0.2–0.4 mm inside the margin of lobes, more or less sunken, appearing prominent on upper surface; indusia cup-shaped, up to 1.2 mm long, 0.7 mm diam.

Distribution in Thailand

EASTERN: Buri Ram; CENTRAL: Nakhon Nayok; PENINSULAR: Surat Thani, Krabi, Nakhon Si Thammarat, Trang, Yala.

Distribution in Laos


Distribution in Cambodia

Kampot, Kompong Speu

Wider Distribution

Burma, S China, Indochina, Hainan, Taiwan, Peninsular Malaysia, Sumatra, Java and New Guinea.


On mossy tree trunks or on rocks near streams in light shade or in clearing and on ridge and at low or medium altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Middleton et al. 4416, Nakhon Si Thammarat, Namtok Yong National Park (E).

Voucher specimens - Laos

Poilane 16152, Salavan, Pou Set (P).

Voucher specimens - Cambodia

Pierre 5770, Kampot, Kamchay (P);Pierre 5773, Kompong Speu (P).







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